Set in Han Imperial China, the plot follows the adventures of enslaved girl Ping with ancient dragon Long Danzi. Dragons had been banished from the kingdom. Ping, an orphan, finds one of the last remaining dragon eggs. Palace guards force Ping to run away in order to return the dragon egg to the ocean and save all dragons from extinction. Ping discovers that she is a true Dragonkeeper.
Ashitaka, a prince of the disappearing Emishi people, is cursed by a demonized boar god and must journey to the west to find a cure. Along the way, he encounters San, a young human woman fighting to protect the forest, and Lady Eboshi, who is trying to destroy it. Ashitaka must find a way to bring balance to this conflict.
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